A new Oriental species of Gnoristinae (Diptera: Mycetophilidae) from Borneo

� Abstract: A new species of Mycetophilidae (Diptera), tentatively placed in Dziedzickia Johannsen, is herein described as D. longistyla sp. nov. (Malaysia: Sarawak). The species described herein differs from the two previously known Oriental species included in Dziedzickia in having simple antenae, laterotergite with long setae and extended, apically pointed gonostyle. A brief discussion of the relationships among some genera of Gnoristinae is also presented.


Introduction
Hitherto, 56 recent species have been placed in Dziedzickia Johannsen, 1909 (Diptera: Mycetophilidae: Gnoristinae): 40 Neotropical, 6 Nearctic, 5 Afrotropical, 3 Palaearctic and 2 Oriental (Bechev, unpublished database), apparently belonging to different genera (Ševčík et al., 2011).The previously described two Oriental species, remarkable with the presence of pectinate antennae, are tentatively placed in the genus (Ševčík et al., 2011).Here we describe a new species, also tentatively placed in Dziedzickia pending a generic revision of the Gnoristinae (see Discussion).The new species differs from the other two Oriental species of Dziedzickia in having simple antenae, laterotergite with long setae and gonostyle narrow, extended, apically pointed, with a relatively clearly visible tooth.

Material and methods
The studied material has been collected by S. Kazandzhieva in Kubah National Park, Borneo (Malaysia: Sarawak) in a mixed dipterocarp forest habitat by Malaise trap and was preserved in ethanol.The habitus photos (specimen in alcohol) were taken by a digital camera Canon EOS 750D with Canon 100 mm Macro lens.The terminalia of the specimen were removed and subsequently macerated in 10% warm KOH.Dissections and temporary slides were made in glycerol.The slides were photographed with the Canon EOS 750D fit to the compound microscope.The terminalia were afterwards transferred to microvial with glycerine and stored together with the specimen.The habitus and male terminalia photos were combined using the Helicon Focus 5 software from multiple gradually Historia naturalis bulgarica 46 (2024) focused images.The morphological terminology follows Søli (2017).
The studied material is deposited in Regional Natural History Museum of Plovdiv, Plovdiv, Bulgaria (RNHMP).
Wing (Figs 2-3).Hyaline unmarked, membrane covered only with microtrichia, without macrotrichia.Costa produced beyond R 4+5 to about 1/3 of the distance between the tips of R 4+5 and M 1 .Sc short, bare, ending in R. R 1 about 0.6 of length of R 4+5 .R-m about 2 times as long as Rs, and 1.4 times as long as stem of M-petiole.Sc-r and R 2+3 absent.Base of Cufork before the base of M-fork.CuP as long as CuApetiole.Setulae present on C, upper side of R, R 1 and R 5 .Other veins bare.Halteres yellow.
Abdomen.Yellow with dark bristles.Tergites 1 to 7 with brown dorsal marks in posterior part, semitriangular in dorsal view and better visible on tergites 1 to 4 (Fig. 2).Sternites yellow, without median sclerotised concave fold.
Terminalia (Figs 4-7).Brown.Tergite 9 short, trapeziform.Cerci placed caudally to tergite 9. Gonocoxites not fused ventrally.Dorsally with apical part rounded, enveloping the base of the gonostylus, and with numerous dark spines and bristles from the inside around the middle.Aedeagus elongate, sclerotized.Gonostylus narrow, extended, apically pointed and with a relatively clearly visible tooth.
Female.Unknown.Etymology.The specific epithet is derived from Latin, longus [long] and stylus [stylus], referring to the narrow, extended gonostylus.
Notes.Mesonotum of the type specimen cracked.Setae on mesonotum and scutellum broken.Right fore and mid legs, left hind leg and fore tarsomeres absent.

Discussion
Being previously pointed out by several authors (Hutson, 1979;Ševčík & Chandler, 2008;Ševčík et al., 2011), a heterogenеous group of species, apparently belonging to different genera, is placed in Dziedzickia sensu lato.The type species of the genus, Dziedzickia marginata Dziedzicki, 1885, is distinguished by the presence of the following combination of characters: mediotergite bare, laterotergite haired, C extending beyond apex of R 5 , Sc ending in R, R 2+3 present, point of furcation of posterior fork distinctly before point of furcation of anterior fork.However, subsequently some authors placed in this genus, also, species with bare laterotergite, Sc ending free or without R 2+3 , apparently belonging to different genera (see Ševčík et al., 2011: 694).
Dziedzickia s. lat. is very similar and shares several characters, mainly of wing venation, with Hadroneura Lundström, 1906 and Palaeodocosia Meunier, 1904.In Hadroneura species with welldocumented male genitalia (kamtshatica Stackelberg, 1943;huron Taber, 2018;martini Ševčík, Hippa & Burdíková, 2021), tergite 9 is densely covered with long setae.In the species of Palaeodocosia, tergite 9 is large, and gonostyle has several lobes (see Hutson et al., 1980;Taber, 2017).Because of the absence of long setae coverage on short tergite 9 and the presence of simple gonostyle, as in some Neotropical Dziedzickia (see Oliveira, 2009), we place the new species in Dziedzickia s. lat.pending a generic