Спасов, Н., Илиев, Н., (in press). Бележки върху животинските костни останки от ранните разкопки (1956—1959) на къснохалколитния обект при Хотница. — В: Ст. Чохаджиев (под редакцията на). Хотница — праисторическа селищна могила. т. I. (2006). Търново.
Spassov, N., Iliev, N., (in press). Analyses of the bones from domestic and wild mammals from the Bronze Age settlement near Baley. — In: St. Alexandrov (ed.). Baley, the settlement from the Late Bronze age. Sofia.
Spassov, N., Georgiev, K., Ivanova, N., Ivanov, V., (in press). Study on the status of the marbled polecat (Vormela peregusna peregusna Guld.) in Southern Dobrudja (Bulgaria). — Bull. of the Mustelidae, Viverridae & Procyonidae group IUCN/SSC.
Spassov, N., Georgiev, K., Ivanov, V., (in press). The Population Status and Dynamics of the Brown Bear (Ursus arctos L.) on the Cross-Border Region between Bulgaria and Greece. — Bull. of the Belgrade Mus. of Nat. Hist.
Спасов, Н., Илиев Н., Боев, З., (in press). Животинските останки от ранно-неолитното селище край с. Капитан-Димитриево ( Пазарджишка област). — В: В. Николов и др. Селищна могила Капитан Димитриево. Разкопки 2001—2005. Под ред. на В. Николов. София, 2011.
Николов, В., Христова, Л., Марков, Г., Богданова, Р., Спасов, Н., 2025. От динозаврите до най-ранните човешки предци — палеонтологията на гръбначните животни в Националния природонаучен музей при БАН. Част 3. — Геология и минерални ресурси, 32 (4—5): 37—43.
Николов, В., Христова, Л., Марков, Г., Богданова, Р., Спасов, Н., 2025. От динозаврите до най-ранните човешки предци — палеонтологията на гръбначните животни в Националния природонаучен музей при БАН. Част 2. — Геология и минерални ресурси, 32 (2—3): 39—44.
Николов, В., Христова, Л., Марков, Г., Богданова, Р., Спасов, Н., 2025. От динозаврите до най-ранните човешки предци — палеонтологията на гръбначните животни в Националния природонаучен музей при БАН. Част 1. — Геология и минерални ресурси, 32 (1): 29—34.
Uzunowa, K., Koychev, B., Peeva, S., Raichev, E., Spassov, N., 2024. Craniometrical sexual dimorphism of the grey wolf (Canis lupus, Canidae, Carnivora) in Bulgaria. — Historia naturalis bulgarica, 46 (5): 127—138.
Tsoukala, E., Nagel, D., Youlatos, D., Crégut-Bonnoure, E., Vlachos, E., Spassov, N., 2024. Primates and carnivores from Late Miocene and earliest Late Pliocene sites of Macedonia, Northern Greece. — Fossil Imprint, 80 (2): 362—389.
Spassov, N., 2024. The Final Pliocene and Early Pleistocene Faunal Dispersals from East to Europe and Correlation of the Villafranchian Biochronology between Eastern and Western Europe. — Quaternary, 7: 43. [PDF]
Milchev, B., Spassov, N., 2024. Invasive alien Coypu (Myocastor coypus) as a prey of the native predator Eurasian Eagle Owl (Bubo bubo) in Bulgaria. — Travaux du Museum National d’Histoire Naturelle “Grigore Antipa”, 67 (2): 443—462. [PDF]
Marciszak, A., Lipecki, G., Spassov, N., 2024. Martes wenzensis Stach, 1959 within the early history of the genus Martes Pinel, 1792. — Acta Geologica Polonica, 74 (1): e4. [PDF]
Gurov, T., Atanassov, E., Yordanov, S., Serbezov, R., Gurova, S.-M., Stanchev, R., Spassov, N., 2024. HPDA Service for Estimating the Brown Bear (Ursus arctos L.) Population in Bulgaria. — In: Dobrinkova, N., Fidanova, S. (eds) Environmental Protection and Disaster Risks (EnviroRISKs 2024). Lecture Notes in Networks and Systems, 883: 104—115. [PDF]
Cazenave, M., Pina, M., Hammond, A. S., Böhme, M., Begun, D. R., Spassov, N., Vecino Gazabon, A., Zanolli, C., Bergeret-Medina, A., Marchi, D., Macchiarelli, R., Wood, B., 2024. Postcranial evidence does not support habitual bipedaism in Sahelanthropus tchadensis: A reply to Daver et al. (2022). — Journal of Human Evolution, 198: 103557. [PDF]
Boev, Z., Nedyalkov, N., Georgiev, D., Spassov, N., 2024. Late Pleistocene birds and mammals from the Kiliite Cave (Central Stara Planina Mts — Central North Bulgaria). — Geologica Balcanica, 53 (1) : 105—115. [PDF]
Baker, K. H., Miller, H., Doherty, S., Gray, H. W., Daujat, J., Çakırlar, C., Spassov, N., Trantalidou, K., Madgwick, R., Lamb, A. L., Ameen, C., Atici, L., Baker, P., Beglane, F., Benkert, H., Bendrey, R., Binois-Roman, A., Carden, R. F., Curci, A., De Cupere, B., Detry, C., Gál, E., Genies, C., Kunst, G. K., Liddiard, R., Nicholson, R. Perdikaris, S., Peters, J., Pigière, F., Pluskowskia, A. G., Sadlerb, P., Sicardc, S., Strid, L., Sudds, J., Symmons, R., Tardio, K., Valenzuelag, A., van Veen, M., Vuković, S., Weinstock, J., Wilkensk, B., Wilson, R. J. A., Evans, J. A., Hoelzel, A. R., Sykes, N., 2024. The 10,000-year biocultural history of fallow deer and its implications for conservation policy. — Proceedings of the National Academy of Sciences, 121 (8): e2310051121. [PDF]
Baker, K. H., Gray, H. W. I., Lister, A. M., Spassov, N., Welch, A. J., Trantalidou, K., De Cupere, B., Bonillas, E., De Jong, M., Çakırlar, C., Sykes, N., Hoelzel, A. R., 2024. Ancient and modern DNA track temporal and spatial population dynamics in the European fallow deer since the Eemian interglacial. — Scientific Reports, 14: 3015. [PDF]
Spassov, N., Ignatov, A., Mihaylov, Ts., 2023. New evidence for the recent presence of the lynx, Lynx lynx (Linnaeus), in Western Stara Planina Mountains, Bulgaria. — Historia naturalis bulgarica, 45: 53—56.
Kampouridis, P., Svorligkou, G., Kargopoulos, N., Spassov, N., Böhme, M., 2023. Revision of the Late Miocene hornless rhinocerotids from Samos Island (Greece) with the designation of neotypes and implications for the European chilotheres. — Journal of Vertebrate Paleontology. [PDF]
Bogdanova, R., Dionisios, Y., Spassov, N., 2023. Terrestriality as reflected in the humerus of Mesopithecus delsoni (Cercopithecidae, Colobinae) from Hadjidimovo, Bulgaria. — Journal of Human Evolution, 180. [PDF]
Nikolov, V., Hristova, L., Spassov, N., 2022. Preliminary data on the metapodial bone histology of hipparionins from the upper Miocene locality Hadjidimovo, Southwestern Bulgaria. — 6th International Symposium on Palaehistology, Programme and Abstracts, 39.
Konstantinov, Y., Spassov, N., Acosta-Pankov, I., 2022. First records of golden jackal and European badger non-antagonistic interaction at the Lower Danube (Bulgaria). — North-Western Journal of Zoology, 18 (1): 100—103.
Spassov, N., Askerov, E., Ignatov, A., Acosta-Pankov, I., 2021. New data on the status of the jungle cat (Felis chaus Schreber, 1777) in Azerbaijan. — Historia naturalis bulgarica, 42: 1—3. doi:10.48027/hnb.42.011
Yaneva, M., Konyovska, R., Garevska, B., Hristova, L., Spassov, N., 2020. Sedimentary study of the Upper Miocene fossiliferous deposits in the Tikvesh Basin, Republic of North Macedonia. — Comptes rendus de l’Académie bulgare des Sciences, 73 (4): 516—520. doi:10.7546/CRABS.2020.04.10
Nikolov, V., Dochev, D., Pavlishina, P., Brusatte, S., Yaneva, M., Konyovska, R., Vergilov, V., Simov, N., Spassov, N., Hristova, L., 2020. Welcome to “Cretaceous Park”: Three years of research at the Late Cretaceous tetrapod fossil site near the town of Tran, Western Srednogorie. — Review of the Bulgarian Geological Society, 81 (3): 141—143.
Karastoyanova, N., Gorczyk, J., Spassov, N., 2020. The natural history of the fallow deer, Dama dama (Linnaeus, 1758) in Bulgaria in prehistory and new evidence for the existence of an autochthonous Holocene population in the Balkans. — Intеrnational Journal of Osteoarchaeology, 30 (5): 616—628. doi:10.1002/oa.2886
Frémondeau, D., Ottoni, C., Ivanova, S., Marinova, E., Spassov, N., Hristova, L., Konyovska, R., Van Neer, W., Lupianez, N, Gurova, M., 2020. New mtDNA and Isotopic Evidence on Late Pleistocene Cave Bears in the Balkans: the Case-study of Magura Cave, NW Bulgaria. — Acta Zoologica Bulgarica, 72 (3): 461—478.
Yankova, I., Marinov, M., Neov, B., Petrova, M., Spassov, N., Hristov, P., Radoslavov, G., 2019. Evidence for Early European Neolithic Dog Dispersal: New Data on Southeastern European Subfossil Dogs from the Prehistoric and Antiquity Ages. — Genes, 10 (10): 757. doi:10.3390/genes10100757
Spassov, N., Geraads, D., Hristova, L., Markov, G. N., 2019. The Late Miocene (Turolian) mammal fauna from Gorna Sushitsa, Southwestern Bulgaria. — Neues Jahrbuch für Geologie und Paläontologie, 291 (3): 317—350.
Spassov, N., Askerov, E., Akosta-Pankov, I., Ignatov, A., 2019. New data on the occurrence of Panthera pardus in the Talysh Mountains, Azerbaijan (Carnivora: Felidae). — Lynx, 50: 113—118.
Spassov, N., Acosta-Pankov, I., 2019. Dispersal history of the golden jackal (Canis aureus moreoticus Geoffroy, 1835) in Europe and possible causes of its recent population explosion. — Biodiversity Data Journal, 7: e34825. doi:10.3897/BDJ.7.e34825
Nishita, Y., Spassov, N., Peeva, S., Raichev, E. G., Kaneko, Y., Masuda, R., 2019. Genetic diversity of MHC class II DRB alleles in the marbled polecat, Vormela peregusna, in Bulgaria. — Ethology Ecology & Evolution, 31 (1): 59—72. doi:10.1080/03949370.2018.1486887
Ersmark, E., Baryshnikov, G., Higham, T., Argant, A., Castaños, P., Döppes, D., Gasparik, M., Germonpré, M., Lidén, K., Lipecki, G., Marciszak, A., Miller, R., Moreno‐García, M., Pacher, M., Robu, M., Rodriguez-Manuel Rojo Guerra, R., Sabol, M., Spassov, N., Storå, J., Valdiosera, C., Villaluenga, A., Stewart, J. R., Dalén, L., 2019. Genetic turnovers and northern survival during the last glacial maximum in European brown bears. — Ecology and Evolution, 9 (10): 5891—5905. doi:10.1002/ece3.5172
Böhme, M., Spassov, N., Fuss, J., Tröscher, A., Deane, A. S., Prieto, J., Kirscher, U., Lechner, T., Begun, D. R., 2019. A new Miocene ape and locomotion in the ancestor of great apes and humans. — Nature. doi:10.1038/s41586-019-1731-0
Boev, Z., Spassov, N., 2019. Past distribution of Castor fiber in Bulgaria: fossil, subfossil and historical records (Rodentia: Castoridae). — Lynx, n. s. (Praha), 50: 37—49. [PDF]
Spassov, N., Iliev, N., Hristova, L., Ivanov, V., 2018. Typology of Thracian horses according to osteological analysis of skeletal remains and depictions from the antiquity. — Historia naturalis bulgarica, 25: 15—24. [PDF] doi:10.5281/zenodo.4043992
Spassov, N., Hristova, L., Iliev, N., 2018. The domesticated horses from the submerged prehistoric village of Urdoviza (Kiten) on the Bulgarian Black Sea coast — among the oldest known. — Historia naturalis bulgarica, 25: 11—14. [PDF] doi:10.5281/zenodo.4043978
Spassov, N., Hristova, L., 2018. Early Ursus etruscus Cuvier from the Middle Villafranchian carst cavern locality of Varshets (Bulgaria). — 24th International Cave Bear Symposium 2018, Chepelare, 27—30.09.2018.
Spassov, N., Geraads, D., Hristova, L., Markov, G. N., Garevska, B., Garevski, R., 2018. The late Miocene mammal faunas of the Republic of Macedonia (FYROM). — Palaeontographica A., 311 (1—6): 1—85.
Mennecart, B., Geraads, D., Spassov, N., Zagorchev, I., 2018. Discovery of the oldest European ruminant in the late Eocene of Bulgaria: Did tectonics influence the diachronic development of the Grande Coupure? — Palaeogeography, Palaeoclimatology, Palaeoecology, 498: 1—8.
Гюрова, М., Иванова, С., Маринова, Е., Попов, В., Спасов, Н., Христова, Л., Ферхейден, С., Бурле, К., 2018. Проучване на пещерата Мишин камик — сезон 5. — Археологически открития и разкопки през 2017 г. БАН. Национален археологически институ с музей. София, с. 8—11.
Fuss, J., Spassov, N., Böhme, M., Begun, D. R., 2018. Response to Benoit and Thackeray ‘A cladistic analysis of Graecopithecus’. — South African Journal of Science, 114 (5/6): Art. #a0267, 2 pp. doi:10.17159/sajs.2018/a0267
Dzhebir, G., Yordanov, G., Yankova, I., Sirakova, D., Petrova, M., Neov, B., Hristov, P., Radoslavov, G., Hristova, L., Spassov, N., 2018. Comparative genetic analysis of subfossil wild horses (from the Neolithic Age and Early Bronze Age) and present-day domestic horses from Bulgaria. — Historia naturalis bulgarica, 25: 3—10. [PDF] doi:10.5281/zenodo.4043974
Dzhebir, G., Yordanov, G., Yankova, I., Sirakova, D., Petrova, M., Neov, B., Hristov, P., Radoslavov, G., Hristova, L., Spassov, N., 2018. Comparative genetic analysis of modern and ancient horses from the Neolithic age in Bulgaria. — Historia naturalis bugarica, 28: 3—10.
Böhme, M., Van Baak, C. G., Prieto, J., Winklhofer, M., Spassov, N., 2018. Late Miocene stratigraphy, palaeoclimate and evolution of the Sandanski Basin (Bulgaria) and the chronology of the Pikermian faunal changes. — Global and planetary change, 170: 1—19.
Acosta-Pankov, I., Spassov, N., Banea, O. C., 2018. Seasonal differences in golden jackals (Canis aureus L. 1758) howling response in Eastern Bulgaria. — Proceedings of the 2nd International Jackal Symposium, Marathon Bay, Attiki Province, Greece, Hell. Zool. Arch., No. 9. ISSN: 1106-2134.
Acosta-Pankov, I., Spassov, N., Banea, O. C., 2018. Seasonal Differences in Howling Response of Golden Jackals Canis aureus L. 1758 (Mammalia: Canidae) in Eastern Bulgaria. — Acta zoologica bulgarica, 70 (4): 539—545.
Acosta-Pankov, I., Ranc, N., Spassov, N., Banea, O. C., 2018. Preliminary results on population density of golden jackals in Bulgaria. — Proceedings of the 2nd International Jackal Symposium, Marathon Bay, Attiki Province, Greece, Hell. Zool. Arch., No. 9. ISSN: 1106-2134.
Spassov, N., Hristova, L., Ivanova, S., Georgiev, I., 2017. First record of the small cave bear in Bulgaria and the taxonomic status of bears of the Ursus savini Andrews — Ursus rossicus Borissiak group. — Fossil imprint, 73 (3—4): 275—291.